Research Article |
Corresponding author: Christophe Praz ( christophe.praz@unine.ch ) Academic editor: Jessica Litman
© 2023 Christophe Praz, Andreas Müller, Dimitri Bénon, Mike Herrmann, Rainer Neumeyer.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Praz C, Müller A, Bénon D, Herrmann M, Neumeyer R (2023) Annotated checklist of the Swiss bees (Hymenoptera, Apoidea, Anthophila): hotspots of diversity in the xeric inner Alpine valleys. Alpine Entomology 7: 219-267. https://doi.org/10.3897/alpento.7.112514
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We present a checklist of the Swiss bees and provide information on the distribution of every bee species in all 26 Swiss cantons. 632 species are reported, including the European honeybee Apis mellifera Linnaeus, 1758 and the exotic species Megachile sculpturalis Smith, 1853. Species richness in each canton was correlated with the canton area, with the four largest cantons hosting the highest number of species. Bee diversity hotspots were located in some Alpine inner valleys characterized by a dry and warm climate due to the rain shadow effects of surrounding mountains. These hotspots are mostly located in the steppe-like habitats of the Valais and Graubünden cantons. They host diverse wild bee communities which include a unique assemblage of submediterranean faunal elements and subalpine species. In addition, these habitats host rare species with strongly disjunct distributions in Europe, further stressing the conservation priority of these habitats for wild bee conservation. Intensive faunistic surveys performed in the last 20 years have revealed that about 20 bee species, either previously unknown for Switzerland or which had disappeared from the country for several decades, have colonised areas close to the borders of France and Italy. Most of these new or reappeared species were observed in the warmest area of the country and presumably colonized or recolonized the country from neighbouring regions following global warming. Lastly, DNA barcodes are presented for 394 specimens, including for many species so far not represented in the BOLD database. The taxonomic status of numerous unclear taxa is briefly discussed based on combined genetic and morphological analyses.
Bees, pollinators, conservation, DNA barcoding, biodiversity, biogeography
Despite its small size, Switzerland has received much attention from entomologists for more than a century. This is particularly true for the Alps which cover nearly 60% of the country, and which have attracted botanists and entomologists for a long time, resulting in numerous studies documenting the Swiss flora and fauna (see for example
The first author who extensively studied the Swiss bee fauna was Emil Frey-Gessner (1826–1917;
Jacques de Beaumont (1901–1985; see
While de Beaumont and Frey-Gessner largely documented the wild bee fauna of western Switzerland, and to a very limited extent that of Ticino and of the National Park region in the Graubünden, large parts of Switzerland remained virtually uninvestigated until about 1960, especially the north-eastern corner, as well as Graubünden with the vast valleys of the upper Rhine river system. Even the eastern part of the Valais, corresponding to the German-speaking part of the Valais, perhaps the most remarkable entomological “hotspot” in Switzerland due to its dry and continental climate, also remained largely unexplored until well after 1950. As noted by
Andrena probata (left) and Andrena ranunculorum (right), two rare and emblematic species of the dry, inner Alpine Valleys of Switzerland. Pictures by Dimitri Bénon (www.swisswildbees.ch).
The intensive work of Erwin Steinmann (1923–2020; see
The Swiss bee fauna is and will forever be associated with the name of Felix Amiet, who has dedicated much of his life to the study of the Swiss bees. Felix Amiet started his work on the bees in the 1960s. After that and until now, he has explored every corner of Switzerland to study its bees. He has assembled the most comprehensive collection to date, with well-preserved specimens from all biogeographic regions of the country. His collection is preserved in the Natural History Museum of Bern. Felix Amiet published the first checklist of the Swiss bees (
The present checklist builds upon the previous work coordinated by Felix Amiet. Since 2012, much faunistic work has been conducted on the Swiss bees in a project aiming at updating the Red List of the Swiss bees; during this project, numerous inventories have been undertaken, partly in poorly investigated regions of Switzerland. Several new species for Switzerland have been found, and our knowledge of the distribution of the bees has substantially improved. In parallel, bee systematic studies have seen a renewed impetus, in part due to the increased use of genetic markers to delineate species, leading to numerous taxonomic changes in the central European fauna. Before the publication of the Swiss red list (Müller and Praz, in press), we wish to summarize all these changes in the present work. We thus present a revised checklist of all Swiss bee species, giving for each species its distribution in all cantons of Switzerland. We also comment on each species: i. if it is new for Switzerland; ii. if it is a recently described species, or a recently recognized taxon previously treated as a synonym; iii. if its current distribution is different from that previously published (
As in other central or northern European countries, the bee fauna of Switzerland has seen massive declines, which gradually began after the main rivers had been channeled and the lowland areas subjected to urbanization and agricultural intensification. The few scientific collections that predate these important changes are thus irreplaceable testimonies of historical bee diversity. Unfortunately, these collections are small and their geographic coverage very fragmentary; moreover, the standards for curating specimens, especially with respect to the locality information on the labels, were not those of today. Consequently, some of these collections are somehow problematic, and some records dubious, for example because the supposed locality information refers to the place of residence of the author, or simply because of labelling mistakes. We followed here the guidelines of
Given the difficulties inherent in the identification of bees, we only validated records that are based on a preserved specimen; some old records based only on literature mentions are not validated (see for example the case of Andrena hystrix Schmiedeknecht, 1883, below). As indicated above, one criterion to validate or reject isolated records is whether the record lies within the overall distribution of the species in Europe (
In some insect groups, occurrences based on isolated individuals are sometimes considered to represent erratic individuals (vagrants) outside their reproductive range. Examples include historical records of butterflies in southern Switzerland, for example Gonepteryx cleopatra (Linnaeus, 1767) in Ticino or Coenonympha dorus (Esper, 1782) in the Geneva region; both species are not considered to be native in Switzerland and were not evaluated in the red list of the Swiss butterflies (
Unlike in some other insect groups, notably the Coleoptera (
The collection of Walter Schmid is slightly more problematic because the specimens were not labeled with original locality data by Schmid himself. All specimens in his collection bear a printed label that simply mentions “Basel, W. Schmid” (Suppl. material
All Swiss bee records, including those based on museum material, faunistic inventories or citizen-science based observations, are assembled into a database centralized at info fauna, Neuchatel (www.infofauna.ch). The main task of info fauna is to maintain faunistic databases, to publish them as open resource at a low resolution (5 × 5 km) nationally and internationally, and to diffuse precise occurrence data to actors active in conservation, such as nature reserve managers and conservation agencies. The entire bee dataset can be accessed freely on GBIF (
Subspecies
Subspecies are not recognized in this dataset. In some cases, the initial database assembled for the publication of the identification keys by
Species aggregates
In bees, there are numerous groups of species where a confident identification is only possible in fresh specimens or when some specific characters are visible, for example the male genitalia. In addition, only one sex can be identified with confidence in several groups. For example, the females of the Halictus simplex group (Halictus simplex Blüthgen, 1923, H. langobardicus Blüthgen, 1944 and H. eurygnathus Blüthgen, 1931) are nearly impossible to separate (
Species aggregate | Included species | Remark |
---|---|---|
Lasioglossum alpigenum/bavaricum aggr. | Lasioglossum alpigenum, L. bavaricum | All females of these two species are included in this species aggregate, unless a DNA-based identification was performed. |
Lasioglossum fratellum/subfulvicorne aggr. | Lasioglossum fratellum, L. subfulvicorne | All females of these two species are included in this species aggregate. |
Hylaeus gibbus aggr. | Hylaeus confusus, H. gibbus, H. incongruus | Most specimens were identified using morphology; specimens that could not be re-rexamined recently, as well as morphologically unclear specimens, were included in this aggregate. |
Panurginus montanus aggr. | Panurginus montanus, P. sericatus | Most females of these two species were included in this aggregate. |
Andrena proxima aggr. | Andrena alutacea, A. ampla, A. proxima | Morphologically unclear specimens (mostly males, but also worn females) were included in this aggregate. All specimens from the Valais upstream from Martigny were assigned to A. ampla ( |
Halictus simplex aggr. | Halictus eurygnathus, H. langobardicus, H. simplex | All females of these three species are included in this species aggregate, unless a DNA-based identification was performed. |
Hylaeus taeniolatus/pictipes aggr. | Hylaeus pictipes, H. taniolatus | Females were systematically assigned to this aggregate until 2019; thereafter, most females were separated based on their morphology. |
Bombus terrestris aggr. | Bombus cryptarum, B. lucorum, B. magnus, B. terrestris | All workers were included in this species aggregate, unless identified using DNA. Queens and males were mostly identified to the species level. |
Generic classification
We adopt here the classification of
For the present work, numerous identifications have been verified using DNA barcoding, following lab protocols published elsewhere (e.g.,
We generated DNA barcodes with the following aims. First, in numerous cases, DNA-assisted identifications allowed to delineate the distribution of some species and generated confidently identified material to evaluate morphological criteria (see for example the case of the Hylaeus gibbus-group, below). Second, we generated DNA barcodes to evaluate the taxonomic status of some “forms” with unclear taxonomy. Lastly, we generated DNA barcodes for Swiss species not yet represented in DNA barcode libraries (www.bold.org, hereafter BOLD). All generated DNA barcodes have been submitted to BOLD; a full list of specimens for which DNA barcodes were generated, including locality information as well as BOLD accession numbers, is given in Suppl. material
The database includes 473653 occurrences between 1817 and 2021 (inclusive), distributed in 7027 1 × 1-km grid cells; all records, even the historical records, are attributed to one 1 × 1-km grid cell, corresponding to the kilometer grid of the national topographic maps (www.map.geo.admin.ch); one occurrence may include several specimens when more than one specimen was collected at the exact same locality on the same day. The evolution of the database in time is shown in Fig.
To date, 632 bee species have been recorded for Switzerland, including one exotic species, Megachile sculpturalis and the European honeybee, Apis mellifera Linnaeus, 1758. The distribution of these species in all Swiss cantons is given in Table
Checklist of the Swiss bees, showing the presence of each species in each canton. Open circles indicate records before 2000, closed circles after 1999. Abbreviations: AG: Aargau; AI: Appenzell Innerrhoden; AR: Appenzell Ausserrhoden; BE: Bern; BL: Basel-Landschaft; BS: Basel-Stadt; FR: Fribourg; GE: Geneva; GL: Glarus; GR: Graubünden; JU: Jura; LU: Luzern; NE: Neuchâtel; NW: Nidwalden; OW: Obwalden; SG: St. Gallen; SH: Schaffhausen; SO: Solothurn; SZ: Schwyz; TG: Thurgau; TI: Ticino; UR: Uri; VD: Vaud; VS: Valais; ZG: Zug. This table is also available as a supplemental table (Suppl. material
Species | AG | AI | AR | BE | BL | BS | FR | GE | GL | GR | JU | LU | NE | NW | OW | SG | SH | SO | SZ | TG | TI | UR | VD | VS | ZG | ZH | Number of Cantons | Number of Cantons (after 2000) |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
Aglaoapis tridentata | ● | ● | ● | ● | ○ | ● | 6 | 5 | ||||||||||||||||||||
Amegilla albigena | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Amegilla garrula | ○ | ● | ● | ○ | 4 | 2 | ||||||||||||||||||||||
Amegilla quadrifasciata | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Amegilla salviae | ○ | 1 | 0 | |||||||||||||||||||||||||
Ammobates punctatus | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena aeneiventris | ● | ○ | ● | ○ | ● | 5 | 3 | |||||||||||||||||||||
Andrena afrensis | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena afzeliella | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Andrena agilissima | ● | ● | ○ | ○ | ● | ● | ● | ● | ○ | ● | ● | ○ | ● | 13 | 9 | |||||||||||||
Andrena alfkenella | ● | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 16 | 14 | ||||||||||
Andrena allosa | ● | ● | ● | ● | ● | ● | 6 | 6 | ||||||||||||||||||||
Andrena alutacea | ○ | ● | ● | ○ | ● | ● | ● | 7 | 5 | |||||||||||||||||||
Andrena amieti | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 11 | 11 | |||||||||||||||
Andrena ampla | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Andrena apicata | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ○ | 17 | 14 | |||||||||
Andrena argentata | ○ | ○ | ○ | ● | ○ | ○ | 6 | 1 | ||||||||||||||||||||
Andrena assimilis | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Andrena barbareae | ● | ● | ● | ○ | ● | 5 | 4 | |||||||||||||||||||||
Andrena barbilabris | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 19 | |||||
Andrena bicolor | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Andrena bimaculata | ○ | ○ | ● | ● | ● | ● | 6 | 4 | ||||||||||||||||||||
Andrena bucephala | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 18 | ||||||||
Andrena chrysopus | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Andrena chrysosceles | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Andrena cineraria | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Andrena clarkella | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | 16 | 14 | ||||||||||
Andrena coitana | ○ | ○ | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | 11 | 7 | |||||||||||||||
Andrena combinata | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 16 | |||||||
Andrena confinis | ○ | ○ | ● | ● | ○ | ○ | ○ | ● | ○ | ○ | 10 | 3 | ||||||||||||||||
Andrena congruens | ○ | ○ | ● | ○ | ○ | ○ | ● | ● | ● | 9 | 4 | |||||||||||||||||
Andrena curvungula | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 15 | |||||||||||
Andrena decipiens | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Andrena denticulata | ● | ○ | ● | ● | ○ | ● | ● | 7 | 5 | |||||||||||||||||||
Andrena distinguenda | ● | ○ | ○ | ● | 4 | 2 | ||||||||||||||||||||||
Andrena dorsata | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Andrena falsifica | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 23 | ||
Andrena ferox | ○ | ○ | ● | ○ | ○ | ○ | ● | 7 | 2 | |||||||||||||||||||
Andrena flavilabris | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena flavipes | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Andrena florea | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 13 | 13 | |||||||||||||
Andrena floricola | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ● | 8 | 1 | ||||||||||||||||||
Andrena florivaga | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena freygessneri | ● | ○ | ○ | ● | 4 | 2 | ||||||||||||||||||||||
Andrena fucata | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 21 | |||
Andrena fulva | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 20 | 19 | ||||||
Andrena fulvago | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Andrena fulvata | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Andrena fulvicornis | ● | ● | ● | ● | 4 | 4 | ||||||||||||||||||||||
Andrena fulvida | ● | ○ | ● | ● | ● | 5 | 4 | |||||||||||||||||||||
Andrena fuscipes | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Andrena fuscosa | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena gelriae | ○ | ○ | ● | ○ | ● | ● | ● | 7 | 4 | |||||||||||||||||||
Andrena gravida | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 22 | ||||
Andrena haemorrhoa | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Andrena hattorfiana | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 23 | |
Andrena helvola | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 23 | 22 | |||
Andrena hesperia | ● | ● | ○ | 3 | 2 | |||||||||||||||||||||||
Andrena humilis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Andrena hypopolia | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Andrena incisa | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Andrena intermedia | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 20 | 17 | ||||||
Andrena labialis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 13 | 13 | |||||||||||||
Andrena labiata | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Andrena lagopus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 17 | 16 | |||||||||
Andrena lapponica | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 14 | |||||||||||
Andrena lathyri | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 19 | ||||||
Andrena lepida | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena limata | ○ | ○ | ○ | ○ | ● | ● | ○ | 7 | 2 | |||||||||||||||||||
Andrena livens | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena marginata | ● | ● | ○ | ● | ● | ○ | ● | ○ | ● | ● | 10 | 7 | ||||||||||||||||
Andrena minutula | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Andrena minutuloides | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Andrena mitis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 17 | |||||||||
Andrena montana | ● | ● | ● | ● | ● | 5 | 5 | |||||||||||||||||||||
Andrena mucida | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Andrena nana | ● | ○ | ○ | ○ | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ● | 17 | 12 | |||||||||
Andrena nanula | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Andrena nigroaenea | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ○ | 21 | 17 | |||||
Andrena nigroolivacea | ● | ● | ● | ● | ● | ● | ● | ● | ● | 9 | 9 | |||||||||||||||||
Andrena nigrospina | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Andrena nitida | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Andrena nitidiuscula | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 17 | 15 | |||||||||
Andrena niveata | ○ | ○ | ○ | ● | ● | 5 | 2 | |||||||||||||||||||||
Andrena nuptialis | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Andrena nycthemera | ○ | ○ | ○ | ○ | ○ | ○ | 6 | 0 | ||||||||||||||||||||
Andrena ovata | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Andrena pallitarsis | ○ | ○ | ○ | ● | ○ | 5 | 1 | |||||||||||||||||||||
Andrena pandellei | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 17 | |||||||||
Andrena parviceps | ○ | ● | ○ | ● | 4 | 2 | ||||||||||||||||||||||
Andrena pauxilla | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena pellucens | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena pilipes | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ● | 8 | 1 | ||||||||||||||||||
Andrena polita | ○ | ● | ● | ● | ● | ● | ● | 7 | 6 | |||||||||||||||||||
Andrena potentillae | ○ | ○ | ○ | ● | ● | ● | 6 | 3 | ||||||||||||||||||||
Andrena praecox | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Andrena probata | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Andrena propinqua | ○ | ○ | ● | ● | ○ | ● | ○ | ● | ● | ○ | 10 | 5 | ||||||||||||||||
Andrena proxima | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Andrena pusilla | ○ | ● | ○ | ○ | 4 | 1 | ||||||||||||||||||||||
Andrena ranunculorum | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena rhenana | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Andrena rogenhoferi | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 12 | 12 | ||||||||||||||
Andrena rosae | ● | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ○ | ● | ● | 20 | 16 | ||||||
Andrena ruficrus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 14 | 14 | ||||||||||||
Andrena rufizona | ○ | ○ | ● | ○ | ● | ● | ● | ○ | 8 | 4 | ||||||||||||||||||
Andrena rufula | ● | ● | ● | ● | ● | ● | 6 | 6 | ||||||||||||||||||||
Andrena rugulosa | ○ | ● | ● | ○ | ● | ● | 6 | 4 | ||||||||||||||||||||
Andrena russula | ○ | ○ | ○ | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 12 | 8 | ||||||||||||||
Andrena saxonica | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena schencki | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | 15 | 11 | |||||||||||
Andrena scotica | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Andrena semilaevis | ● | ○ | ○ | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 13 | ||||||||||
Andrena sericata | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Andrena simillima | ● | ○ | ● | 3 | 2 | |||||||||||||||||||||||
Andrena simontornyella | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Andrena strohmella | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Andrena subopaca | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Andrena suerinensis | ● | ○ | 2 | 1 | ||||||||||||||||||||||||
Andrena symphyti | ● | ○ | 2 | 1 | ||||||||||||||||||||||||
Andrena synadelpha | ○ | ● | ○ | ○ | ○ | ● | ● | 7 | 3 | |||||||||||||||||||
Andrena taraxaci | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Andrena tarsata | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | 10 | 8 | ||||||||||||||||
Andrena tenuistriata | ● | 1 | 1 | |||||||||||||||||||||||||
Andrena thoracica | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Andrena tibialis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 19 | |||||||
Andrena tridentata | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena trimmerana | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 15 | 12 | |||||||||||
Andrena tscheki | ○ | 1 | 0 | |||||||||||||||||||||||||
Andrena vaga | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Andrena varians | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ● | ● | 12 | 2 | ||||||||||||||
Andrena ventralis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 20 | |||||
Andrena viridescens | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Andrena vulpecula | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Andrena wilkella | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Anthidiellum strigatum | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 23 | ||
Anthidium cingulatum | ○ | ○ | ● | ○ | ○ | ○ | 6 | 1 | ||||||||||||||||||||
Anthidium florentinum | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Anthidium loti | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Anthidium manicatum | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Anthidium montanum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Anthidium oblongatum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Anthidium punctatum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Anthidium septemspinosum | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Anthophora aestivalis | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 20 | |||||
Anthophora balneorum | ○ | ○ | ● | 3 | 1 | |||||||||||||||||||||||
Anthophora bimaculata | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Anthophora canescens | ○ | 1 | 0 | |||||||||||||||||||||||||
Anthophora crassipes | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Anthophora crinipes | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 10 | 9 | ||||||||||||||||
Anthophora dispar | ● | 1 | 1 | |||||||||||||||||||||||||
Anthophora fulvitarsis | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Anthophora furcata | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Anthophora mucida | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Anthophora plagiata | ● | ○ | ○ | ● | ○ | ● | ● | 7 | 4 | |||||||||||||||||||
Anthophora plumipes | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Anthophora pubescens | ○ | ● | ○ | ● | ● | 5 | 3 | |||||||||||||||||||||
Anthophora quadrimaculata | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Anthophora retusa | ● | ● | ● | ● | 4 | 4 | ||||||||||||||||||||||
Apis mellifera | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Biastes emarginatus | ○ | ● | ● | 3 | 2 | |||||||||||||||||||||||
Biastes truncatus | ● | ○ | 2 | 1 | ||||||||||||||||||||||||
Bombus alpinus | ● | ● | ○ | ● | ● | ● | ● | 7 | 6 | |||||||||||||||||||
Bombus argillaceus | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Bombus barbutellus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus bohemicus | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Bombus campestris | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Bombus confusus | ○ | ○ | ○ | ● | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | 12 | 1 | ||||||||||||||
Bombus cryptarum | ● | ○ | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | 12 | 9 | ||||||||||||||
Bombus distinguendus | ○ | ○ | ○ | ● | 4 | 1 | ||||||||||||||||||||||
Bombus flavidus | ● | ● | ● | ● | ○ | ● | ○ | ● | ● | 9 | 7 | |||||||||||||||||
Bombus gerstaeckeri | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 14 | 14 | ||||||||||||
Bombus hortorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus humilis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Bombus hypnorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus inexspectatus | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Bombus jonellus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Bombus lapidarius | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus lucorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus magnus | ○ | 1 | 0 | |||||||||||||||||||||||||
Bombus mendax | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 14 | |||||||||||
Bombus mesomelas | ○ | ● | ● | ○ | ● | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | 16 | 12 | ||||||||||
Bombus monticola | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 14 | |||||||||||
Bombus mucidus | ● | ● | ● | ○ | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 14 | ||||||||||
Bombus muscorum | ● | ○ | ○ | ● | ○ | ● | ○ | ● | ○ | ● | ● | ○ | ● | ○ | ● | ● | 16 | 9 | ||||||||||
Bombus norvegicus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Bombus pascuorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus pomorum | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | 16 | 0 | ||||||||||
Bombus pratorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus pyrenaeus | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 14 | 13 | ||||||||||||
Bombus quadricolor | ● | ● | ● | ○ | ● | ● | ○ | ○ | ○ | ○ | ○ | ● | ○ | ● | ● | ● | ● | ○ | 18 | 10 | ||||||||
Bombus ruderarius | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 24 | 22 | ||
Bombus ruderatus | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 17 | ||||||||
Bombus rupestris | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Bombus sichelii | ○ | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 17 | 13 | |||||||||
Bombus soroeensis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Bombus subterraneus | ● | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 16 | |||||||
Bombus sylvarum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 23 | 21 | |||
Bombus sylvestris | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Bombus terrestris | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Bombus vestalis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 22 | ||||
Bombus veteranus | ● | ● | ○ | ● | ○ | ○ | ● | ● | ● | ○ | ○ | ● | ○ | ● | ● | ○ | ● | ● | 18 | 11 | ||||||||
Bombus wurflenii | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 24 | 22 | ||
Ceratina chalcites | ● | 1 | 1 | |||||||||||||||||||||||||
Ceratina chalybea | ● | ● | ● | ○ | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 13 | 11 | |||||||||||||
Ceratina cucurbitina | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 15 | |||||||||||
Ceratina cyanea | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 23 | ||
Ceratina gravidula | ● | ● | ○ | 3 | 2 | |||||||||||||||||||||||
Ceratina nigrolabiata | ● | 1 | 1 | |||||||||||||||||||||||||
Chelostoma campanularum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Chelostoma distinctum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 20 | ||||||
Chelostoma emarginatum | ○ | ● | ○ | 3 | 1 | |||||||||||||||||||||||
Chelostoma florisomne | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Chelostoma foveolatum | ● | ○ | ● | ● | 4 | 3 | ||||||||||||||||||||||
Chelostoma grande | ● | ○ | ○ | ○ | ● | ● | ● | 7 | 4 | |||||||||||||||||||
Chelostoma rapunculi | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Coelioxys afer | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 15 | |||||||||||
Coelioxys alatus | ○ | ● | ○ | ● | ● | 5 | 3 | |||||||||||||||||||||
Coelioxys aurolimbatus | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 18 | |||||||
Coelioxys conicus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 17 | ||||||||
Coelioxys conoideus | ● | ○ | ○ | ● | ○ | ● | ● | ○ | ○ | ○ | ● | ● | ○ | 13 | 6 | |||||||||||||
Coelioxys echinatus | ● | ● | ● | ● | ● | ● | 6 | 6 | ||||||||||||||||||||
Coelioxys elongatus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 19 | |||||||
Coelioxys emarginatus | ○ | 1 | 0 | |||||||||||||||||||||||||
Coelioxys inermis | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Coelioxys lanceolatus | ● | ● | ● | ○ | ● | ● | ○ | ● | 8 | 6 | ||||||||||||||||||
Coelioxys mandibularis | ● | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ○ | ● | ● | 13 | 10 | |||||||||||||
Coelioxys rufescens | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Colletes collaris | ○ | 1 | 0 | |||||||||||||||||||||||||
Colletes cunicularius | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 22 | ||||
Colletes daviesanus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 18 | ||||||||
Colletes floralis | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Colletes fodiens | ○ | ○ | ○ | ● | ● | 5 | 2 | |||||||||||||||||||||
Colletes gallicus | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Colletes hederae | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 22 | ||||
Colletes impunctatus | ● | ● | ● | ● | ● | ● | 6 | 6 | ||||||||||||||||||||
Colletes marginatus | ○ | ○ | ● | ● | 4 | 2 | ||||||||||||||||||||||
Colletes mlokossewiczi | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Colletes nigricans | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Colletes sierrensis | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Colletes similis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Colletes succinctus | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Dasypoda argentata | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Dasypoda hirtipes | ● | ● | ● | ● | ● | ● | ○ | ● | ○ | ● | ○ | 11 | 8 | |||||||||||||||
Dioxys cinctus | ● | ○ | ● | ○ | ○ | ○ | ○ | ● | ○ | 9 | 3 | |||||||||||||||||
Dufourea alpina | ● | ● | ● | ● | ● | ● | ● | ○ | ● | 9 | 8 | |||||||||||||||||
Dufourea dentiventris | ● | ○ | ● | ● | ● | ○ | ○ | ○ | ● | ● | ● | ● | ● | 13 | 9 | |||||||||||||
Dufourea halictula | ● | 1 | 1 | |||||||||||||||||||||||||
Dufourea inermis | ○ | ● | ○ | ● | ● | 5 | 3 | |||||||||||||||||||||
Dufourea minuta | ○ | ● | ○ | ● | ● | 5 | 3 | |||||||||||||||||||||
Dufourea paradoxa | ● | ● | ○ | ● | ● | ● | 6 | 5 | ||||||||||||||||||||
Epeoloides coecutiens | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | 17 | 15 | |||||||||
Epeolus alpinus | ● | ○ | ● | ○ | ○ | ● | 6 | 3 | ||||||||||||||||||||
Epeolus cruciger | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 13 | 12 | |||||||||||||
Epeolus fallax | ● | 1 | 1 | |||||||||||||||||||||||||
Epeolus productulus | ● | 1 | 1 | |||||||||||||||||||||||||
Epeolus variegatus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 18 | ||||||||
Eucera hungarica | ○ | 1 | 0 | |||||||||||||||||||||||||
Eucera interrupta | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Eucera longicornis | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Eucera nigrescens | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Eucera nigrifacies | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Eucera pollinosa | ● | ● | ○ | 3 | 2 | |||||||||||||||||||||||
Halictus carinthiacus | ● | ● | ● | ● | ● | 5 | 5 | |||||||||||||||||||||
Halictus confusus | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 16 | |||||||||
Halictus eurygnathus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | 12 | 11 | ||||||||||||||
Halictus fulvipes | ○ | 1 | 0 | |||||||||||||||||||||||||
Halictus kessleri | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Halictus langobardicus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 15 | 14 | |||||||||||
Halictus leucaheneus | ● | ● | ● | ● | ○ | 5 | 4 | |||||||||||||||||||||
Halictus maculatus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 20 | 19 | ||||||
Halictus quadricinctus | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | 10 | 9 | ||||||||||||||||
Halictus rubicundus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Halictus scabiosae | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Halictus seladonius | ○ | ● | ● | 3 | 2 | |||||||||||||||||||||||
Halictus sexcinctus | ○ | ● | ● | ○ | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | 14 | 10 | ||||||||||||
Halictus simplex | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Halictus subauratus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Halictus submediterraneus | ● | ● | ○ | ● | ● | 5 | 4 | |||||||||||||||||||||
Halictus tectus | ○ | 1 | 0 | |||||||||||||||||||||||||
Halictus tumulorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Heriades crenulata | ● | ○ | ● | ● | ○ | ● | ● | ○ | ● | ● | 10 | 7 | ||||||||||||||||
Heriades rubicola | ● | ● | ○ | 3 | 2 | |||||||||||||||||||||||
Heriades truncorum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Hoplitis acuticornis | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Hoplitis adunca | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Hoplitis anthocopoides | ○ | ○ | ○ | ○ | ○ | ○ | ● | ● | ○ | 9 | 2 | |||||||||||||||||
Hoplitis claviventris | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Hoplitis dalmatica | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Hoplitis lepeletieri | ● | ● | ○ | ● | ○ | ○ | ● | ● | ● | ● | 10 | 7 | ||||||||||||||||
Hoplitis leucomelana | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Hoplitis loti | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 14 | 13 | ||||||||||||
Hoplitis mitis | ● | ● | ○ | ○ | ● | ● | ○ | ● | ● | ○ | ● | ○ | ● | ● | ● | 15 | 10 | |||||||||||
Hoplitis papaveris | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Hoplitis praestans | ● | 1 | 1 | |||||||||||||||||||||||||
Hoplitis ravouxi | ● | ● | ● | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ○ | 14 | 11 | ||||||||||||
Hoplitis robusta | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Hoplitis stecki | ○ | 1 | 0 | |||||||||||||||||||||||||
Hoplitis tridentata | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Hoplitis tuberculata | ● | ○ | ● | ○ | ● | ● | ● | ○ | ○ | ● | ○ | ○ | ● | ● | ● | ● | 16 | 10 | ||||||||||
Hoplitis villosa | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 16 | |||||||||
Hylaeus alpinus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ○ | 12 | 10 | ||||||||||||||
Hylaeus angustatus | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ○ | ● | ○ | ○ | ● | ○ | 15 | 10 | |||||||||||
Hylaeus annulatus | ● | ● | ● | ● | ● | ● | ● | ○ | ● | 9 | 8 | |||||||||||||||||
Hylaeus bifasciatus | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Hylaeus brevicornis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Hylaeus cardioscapus | ● | 1 | 1 | |||||||||||||||||||||||||
Hylaeus clypearis | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 16 | |||||||||
Hylaeus communis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Hylaeus confusus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Hylaeus cornutus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 15 | |||||||||||
Hylaeus crassanus | ○ | ○ | ● | 3 | 1 | |||||||||||||||||||||||
Hylaeus difformis | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 21 | ||||
Hylaeus dilatatus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 19 | |||||||
Hylaeus duckei | ● | ○ | ○ | ● | ● | ○ | ○ | ● | 8 | 4 | ||||||||||||||||||
Hylaeus euryscapus | ○ | 1 | 0 | |||||||||||||||||||||||||
Hylaeus gibbus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Hylaeus glacialis | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Hylaeus gredleri | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 22 | 22 | ||||
Hylaeus hyalinatus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Hylaeus incongruus | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Hylaeus intermedius | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Hylaeus kahri | ● | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 15 | |||||||||
Hylaeus leptocephalus | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | 15 | 11 | |||||||||||
Hylaeus moricei | ○ | ● | ● | 3 | 2 | |||||||||||||||||||||||
Hylaeus nigritus | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Hylaeus nivalis | ● | ● | ● | ● | ● | ● | ● | ● | 8 | 8 | ||||||||||||||||||
Hylaeus paulus | ● | ● | ○ | ● | ● | ● | ○ | ● | ● | ○ | ● | 11 | 8 | |||||||||||||||
Hylaeus pectoralis | ● | ● | ○ | ● | ○ | ● | 6 | 4 | ||||||||||||||||||||
Hylaeus pfankuchi | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 13 | 12 | |||||||||||||
Hylaeus pictipes | ● | ● | ● | ○ | ● | ○ | ○ | ● | ● | 9 | 6 | |||||||||||||||||
Hylaeus pilosulus | ● | 1 | 1 | |||||||||||||||||||||||||
Hylaeus punctatus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Hylaeus punctulatissimus | ● | ● | ○ | ○ | ○ | ○ | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | 15 | 9 | |||||||||||
Hylaeus rinki | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ○ | ● | ● | 17 | 14 | |||||||||
Hylaeus signatus | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 20 | |||||
Hylaeus sinuatus | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Hylaeus styriacus | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 20 | |||||
Hylaeus taeniolatus | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | 17 | 15 | |||||||||
Hylaeus tyrolensis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ○ | 13 | 11 | |||||||||||||
Hylaeus variegatus | ● | ○ | ○ | ● | ● | ● | ○ | ○ | ○ | ● | ● | 11 | 6 | |||||||||||||||
Icteranthidium laterale | ○ | ○ | ● | 3 | 1 | |||||||||||||||||||||||
Lasioglossum aeratum | ● | ● | ○ | ○ | ● | ○ | ● | ● | ● | 9 | 6 | |||||||||||||||||
Lasioglossum albipes | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 25 |
Lasioglossum albocinctum | ● | ○ | ● | 3 | 2 | |||||||||||||||||||||||
Lasioglossum alpigenum | ● | ● | ● | ● | ● | ● | ● | ○ | ○ | ● | ● | 11 | 9 | |||||||||||||||
Lasioglossum angusticeps | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Lasioglossum bavaricum | ● | ● | ● | ● | ● | ○ | ● | ● | ● | 9 | 8 | |||||||||||||||||
Lasioglossum bluethgeni | ● | ● | ● | ● | ● | ● | ● | ● | 8 | 8 | ||||||||||||||||||
Lasioglossum brevicorne | ○ | ● | ● | ○ | ● | 5 | 3 | |||||||||||||||||||||
Lasioglossum breviventre | ○ | ○ | ● | 3 | 1 | |||||||||||||||||||||||
Lasioglossum buccale | ○ | ● | ● | 3 | 2 | |||||||||||||||||||||||
Lasioglossum calceatum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Lasioglossum clypeare | ○ | ● | ○ | ○ | ● | 5 | 2 | |||||||||||||||||||||
Lasioglossum convexiusculum | ○ | ○ | ○ | ● | ○ | ● | ● | 7 | 3 | |||||||||||||||||||
Lasioglossum costulatum | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 13 | |||||||||||
Lasioglossum cupromicans | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 16 | ||||||||||
Lasioglossum discum | ● | ● | ○ | 3 | 2 | |||||||||||||||||||||||
Lasioglossum elegans | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Lasioglossum euboeense | ○ | ○ | ● | 3 | 1 | |||||||||||||||||||||||
Lasioglossum fratellum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 15 | 15 | |||||||||||
Lasioglossum fulvicorne | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Lasioglossum glabriusculum | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 18 | |||||||
Lasioglossum griseolum | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Lasioglossum intermedium | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 17 | ||||||||
Lasioglossum interruptum | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 19 | 17 | |||||||
Lasioglossum laeve | ○ | ○ | ○ | ○ | ○ | ● | 6 | 1 | ||||||||||||||||||||
Lasioglossum laevidorsum | ● | ● | ● | ○ | 4 | 3 | ||||||||||||||||||||||
Lasioglossum laevigatum | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 25 | 21 | |
Lasioglossum laterale | ● | 1 | 1 | |||||||||||||||||||||||||
Lasioglossum laticeps | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Lasioglossum lativentre | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Lasioglossum leucopus | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | 24 | 21 | ||
Lasioglossum leucozonium | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Lasioglossum limbellum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | 18 | 16 | ||||||||
Lasioglossum lineare | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 15 | ||||||||||
Lasioglossum lissonotum | ○ | ○ | ● | ○ | ○ | ● | 6 | 2 | ||||||||||||||||||||
Lasioglossum lucidulum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 16 | ||||||||||
Lasioglossum majus | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | 11 | 10 | |||||||||||||||
Lasioglossum malachurum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Lasioglossum marginatum | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 13 | 12 | |||||||||||||
Lasioglossum marginellum | ● | ○ | ● | 3 | 2 | |||||||||||||||||||||||
Lasioglossum medinai | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Lasioglossum minutissimum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 15 | 14 | |||||||||||
Lasioglossum minutulum | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | 11 | 9 | |||||||||||||||
Lasioglossum monstrificum | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 15 | 14 | |||||||||||
Lasioglossum morio | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Lasioglossum nigripes | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 20 | |||||
Lasioglossum nitidiusculum | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 18 | ||||||
Lasioglossum nitidulum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 24 | 23 | ||
Lasioglossum pallens | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 16 | ||||||||||
Lasioglossum parvulum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Lasioglossum pauperatum | ○ | ○ | ○ | ○ | 4 | 0 | ||||||||||||||||||||||
Lasioglossum pauxillum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Lasioglossum peregrinum | ○ | 1 | 0 | |||||||||||||||||||||||||
Lasioglossum pleurospeculum | ○ | ● | ● | ● | ● | 5 | 4 | |||||||||||||||||||||
Lasioglossum podolicum | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Lasioglossum politum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 20 | ||||||
Lasioglossum punctatissimum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Lasioglossum puncticolle | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | 11 | 10 | |||||||||||||||
Lasioglossum pygmaeum | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 12 | 10 | ||||||||||||||
Lasioglossum quadrinotatulum | ○ | ● | ● | ● | ● | 5 | 4 | |||||||||||||||||||||
Lasioglossum quadrinotatum | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Lasioglossum quadrisignatum | ○ | ○ | ○ | ○ | ● | 5 | 1 | |||||||||||||||||||||
Lasioglossum rufitarse | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ○ | ● | 23 | 21 | |||
Lasioglossum semilucens | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 20 | ||||||
Lasioglossum setulosum | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Lasioglossum sexmaculatum | ○ | 1 | 0 | |||||||||||||||||||||||||
Lasioglossum sexnotatum | ○ | ● | ● | ○ | ○ | ○ | ● | ● | ○ | ○ | ● | ○ | ● | ○ | ● | ○ | 16 | 7 | ||||||||||
Lasioglossum sexstrigatum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 17 | |||||||||
Lasioglossum subfasciatum | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ● | ● | ○ | 12 | 2 | ||||||||||||||
Lasioglossum subfulvicorne | ● | ● | ● | ● | ● | ● | 6 | 6 | ||||||||||||||||||||
Lasioglossum subhirtum | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Lasioglossum tarsatum | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Lasioglossum transitorium | ● | 1 | 1 | |||||||||||||||||||||||||
Lasioglossum tricinctum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 12 | 12 | ||||||||||||||
Lasioglossum villosulum | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Lasioglossum xanthopus | ● | ● | ● | ○ | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 13 | ||||||||||
Lasioglossum zonulum | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 23 | ||
Lithurgus chrysurus | ● | ○ | ● | 3 | 2 | |||||||||||||||||||||||
Macropis europaea | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Macropis fulvipes | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 19 | |||||||
Megachile alpicola | ● | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 22 | ||
Megachile analis | ● | ● | ○ | ● | ● | ● | 6 | 5 | ||||||||||||||||||||
Megachile apicalis | ● | ● | ○ | ○ | ○ | ○ | 6 | 2 | ||||||||||||||||||||
Megachile argentata | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 19 | ||||||
Megachile centuncularis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Megachile circumcincta | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 23 | |||
Megachile ericetorum | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 23 | |
Megachile flabellipes | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Megachile genalis | ● | 1 | 1 | |||||||||||||||||||||||||
Megachile lagopoda | ○ | ○ | ○ | ● | ○ | ● | ● | ● | ○ | 9 | 4 | |||||||||||||||||
Megachile lapponica | ● | ● | ● | ● | ○ | 5 | 4 | |||||||||||||||||||||
Megachile leachella | ○ | ● | ● | ● | 4 | 3 | ||||||||||||||||||||||
Megachile ligniseca | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 20 | |||||
Megachile maritima | ● | ● | ○ | ○ | ● | ○ | ● | ● | ● | ○ | ● | ● | ● | ○ | 14 | 9 | ||||||||||||
Megachile melanopyga | ● | ● | ● | 3 | 3 | |||||||||||||||||||||||
Megachile nigriventris | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | 25 | 24 | |
Megachile parietina | ● | ○ | ○ | ○ | ○ | ○ | ● | ● | ○ | ● | ● | ○ | ● | ● | ○ | ● | ● | 17 | 9 | |||||||||
Megachile pilicrus | ● | ● | ○ | 3 | 2 | |||||||||||||||||||||||
Megachile pyrenaea | ● | ○ | ● | ● | ○ | ○ | ● | ● | ● | ● | 10 | 7 | ||||||||||||||||
Megachile pyrenaica | ● | ○ | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | 12 | 9 | ||||||||||||||
Megachile rotundata | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 12 | 12 | ||||||||||||||
Megachile sculpturalis | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 11 | 11 | |||||||||||||||
Megachile versicolor | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 22 | 20 | ||||
Megachile willughbiella | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Melecta albifrons | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 21 | 21 | |||||
Melecta festiva | ● | 1 | 1 | |||||||||||||||||||||||||
Melecta luctuosa | ○ | ○ | ○ | ○ | ● | ○ | ● | ○ | ● | ● | ● | ● | 12 | 6 | ||||||||||||||
Melitta dimidiata | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Melitta haemorrhoidalis | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 23 | 22 | |||
Melitta leporina | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 18 | ||||||||
Melitta nigricans | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 20 | 19 | ||||||
Melitta tricincta | ● | ○ | ● | ● | ● | ● | ● | ● | 8 | 7 | ||||||||||||||||||
Melitturga clavicornis | ● | 1 | 1 | |||||||||||||||||||||||||
Nomada alboguttata | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 17 | |||||||
Nomada argentata | ○ | ● | ● | ○ | ○ | ● | 6 | 3 | ||||||||||||||||||||
Nomada armata | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 17 | 14 | |||||||||
Nomada atroscutellaris | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 17 | ||||||||
Nomada baccata | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Nomada bifasciata | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | 21 | 19 | |||||
Nomada bispinosa | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Nomada blepharipes | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Nomada bluethgeni | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada braunsiana | ○ | ○ | ○ | ● | ○ | ○ | ● | ● | 8 | 3 | ||||||||||||||||||
Nomada carnifex | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada castellana | ● | ○ | ● | ○ | ○ | ● | ● | ○ | ○ | ● | ● | ● | ○ | ● | ● | ● | 16 | 10 | ||||||||||
Nomada confinis | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada conjungens | ● | ● | ● | ○ | ○ | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ○ | ● | ● | ○ | 18 | 12 | ||||||||
Nomada connectens | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada discrepans | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada distinguenda | ● | ○ | ● | ● | ○ | ● | ○ | ● | ● | ● | ○ | ○ | ○ | ● | 14 | 8 | ||||||||||||
Nomada emarginata | ○ | ○ | ● | ● | ○ | ● | ● | ● | 8 | 5 | ||||||||||||||||||
Nomada errans | ○ | ○ | ● | ● | ● | ○ | ○ | ○ | 8 | 3 | ||||||||||||||||||
Nomada fabriciana | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 25 | 25 | |
Nomada facilis | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | 19 | 16 | |||||||
Nomada femoralis | ○ | ○ | ● | ○ | ● | ○ | ● | ○ | ● | ○ | 10 | 4 | ||||||||||||||||
Nomada ferruginata | ● | ● | ○ | ● | ● | ● | ● | ○ | ○ | ● | ● | ○ | 12 | 8 | ||||||||||||||
Nomada flava | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Nomada flavilabris | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada flavoguttata | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 26 | 26 |
Nomada flavopicta | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 17 | ||||||||
Nomada fucata | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 18 | 17 | ||||||||
Nomada fulvicornis | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 17 | |||||||
Nomada furva | ○ | ○ | ● | 3 | 1 | |||||||||||||||||||||||
Nomada fuscicornis | ○ | ● | ● | ● | ○ | 5 | 3 | |||||||||||||||||||||
Nomada goodeniana | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 23 | ||
Nomada gransassoi | ● | ○ | ○ | ● | 4 | 2 | ||||||||||||||||||||||
Nomada guttulata | ○ | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 16 | 13 | ||||||||||
Nomada hirtipes | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 17 | 16 | |||||||||
Nomada integra | ● | ● | ● | ● | ○ | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 22 | ||
Nomada italica | ○ | 1 | 0 | |||||||||||||||||||||||||
Nomada kohli | ● | ● | ● | ● | 4 | 4 | ||||||||||||||||||||||
Nomada lathburiana | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ● | 20 | 18 | ||||||
Nomada leucophthalma | ● | ○ | ● | ○ | ● | ● | ○ | ● | ● | ○ | ● | ○ | ○ | ● | ● | ● | 16 | 10 | ||||||||||
Nomada linsenmaieri | ● | 1 | 1 | |||||||||||||||||||||||||
Nomada marshamella | ● | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | 25 | 23 | |
Nomada mauritanica | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Nomada melathoracica | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ○ | ● | ○ | 10 | 1 | ||||||||||||||||
Nomada minuscula | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Nomada moeschleri | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 13 | 13 | |||||||||||||
Nomada mutabilis | ○ | ● | 2 | 1 | ||||||||||||||||||||||||
Nomada mutica | ○ | ○ | ● | ○ | ● | ● | ● | 7 | 4 | |||||||||||||||||||
Nomada nobilis | ● | 1 | 1 | |||||||||||||||||||||||||
Nomada obscura | ● | ● | ● | ○ | ● | ● | 6 | 5 | ||||||||||||||||||||
Nomada obtusifrons | ● | ● | ○ | ● | 4 | 3 | ||||||||||||||||||||||
Nomada opaca | ○ | ○ | ○ | ○ | ○ | 5 | 0 | |||||||||||||||||||||
Nomada panurgina | ○ | ○ | 2 | 0 | ||||||||||||||||||||||||
Nomada panzeri | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 24 | ||
Nomada piccioliana | ○ | ● | ○ | ● | ● | ○ | ○ | 7 | 3 | |||||||||||||||||||
Nomada pleurosticta | ● | ● | 2 | 2 | ||||||||||||||||||||||||
Nomada posthuma | ● | ● | ● | ○ | ● | ● | ● | 7 | 6 | |||||||||||||||||||
Nomada rhenana | ○ | ○ | ● | ○ | ○ | ○ | 6 | 1 | ||||||||||||||||||||
Nomada roberjeotiana | ○ | ○ | ○ | 3 | 0 | |||||||||||||||||||||||
Nomada ruficornis | ● | ● | ● | ● | ○ | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 24 | 22 | ||
Nomada rufipes | ○ | ○ | ● | ● | ● | ● | 6 | 4 | ||||||||||||||||||||
Nomada sexfasciata | ● | ● | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 20 | 19 | ||||||
Nomada sheppardana | ● | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ○ | ● | ● | ● | ○ | ● | 16 | 13 | ||||||||||
Nomada signata | ● | ● | ● | ● | ● | ● | ● | ○ | ○ | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | 19 | 17 | |||||||
Nomada similis | ○ | ● | ● | ○ | ● | 5 | 3 | |||||||||||||||||||||
Nomada stigma | ○ | ○ | ○ | ● | ● | ○ | ○ | ○ | ○ | ○ | 10 | 2 | ||||||||||||||||
Nomada striata | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● | ● |