Phylogenetic tree based on maximum likelihood analyses of sequence data of the mitochondrial gene COI and of the three nuclear genes conserved ATPase domain (CAD), elongation factor 1-alpha (EF) and long-wavelength rhodopsin (Opsin). Numbers above branches: statistical support based on 1000 bootstrap replicates (values below 50 are omitted) in the analysis with 4 partitions (by gene); number below branches: support in analysis with 7 partitions (by codon position; a hyphen under a node indicates that this node was not recovered in the analysis); the topology is from the analysis with 4 partitions.